Ed Miliband has clashed with SNP
leader Nicola Sturgeon after she said she would only support him as PM
if he showed he was "better than the Tories".
She said she wanted a "progressive" alliance with Plaid Cymru's Leanne Wood and the Green Party's Natalie Bennett.Mr Miliband, Labour leader, said he wanted to win a majority and would not form a coalition with the SNP.
UKIP leader Nigel Farage also clashed with Mr Miliband on the economy and immigration in the BBC election debate.
- You can watch the debate reaction and follow text commentary here
During the 90-minute programme the Labour leader faced repeated calls from Nicola Sturgeon and Leanne Wood to reject austerity and form a "progressive" alliance with their parties to keep the Conservatives out of government.
Ms Sturgeon claimed he was "so scared to be bold" that he was "not even doing the right thing by the NHS", telling the audience that "if Labour won't be bold enough on its own, I think people should vote for parties that will hold Labour to account and make them bolder".
Ms Sturgeon said this election was about "seizing an alternative to austerity" and asked Mr Miliband whether he would really prefer David Cameron as prime minister than to work with the SNP.
"The difference is I have fought Tories all my life," said Mr Miliband, a reference to the SNP's reliance on Conservative support when they were a minority government at Holyrood.
'Astonishing'
Nigel Farage set his stall out early as being against the "politically correct" consensus - but he riled the audience at London's Methodist Central Hall, saying they were remarkable "even by the left-wing standards of the BBC".Amid boos from the hall, host David Dimbleby said the audience had been selected by a polling company, not the corporation.
"Just astonishing," said Mr Farage in response to the SNP leader's assertion that not all problems in the UK are caused by immigration.
"Yes you are," replied Ms Sturgeon.
'Real alternative'
Mr Farage also repeated his claim that the NHS was spending too much on treating foreign nationals with HIV, saying most people in Britain would agree with him and the NHS should not be an "international health service".The UKIP leader was accused by Ms Bennett of "demonising" migrants and by Ms Sturgeon of "intolerance". Mr Miliband also accused him of "exploiting people's fears".
He then claimed Mr Farage's real agenda was to get rid of the NHS and replace it with a private health insurance - which sparked a furious reaction from the UKIP leader, who repeatedly accused Mr Miliband of "lying" about his beliefs.
Plaid Cymru leader Leanne Wood challenged Mr Miliband to hold an "emergency budget" to end austerity if he becomes prime minister, but Mr Miliband did not rise to the bait, insisting Labour had plans that would help working families.
In his opening statement Mr Farage had said UKIP could cut taxes as part of a fully-costed economic plan and was the only party "prepared to talk straight" and not afraid to upset the "politically correct" consensus.
Leanne Wood was the first party leader to make an opening statement, telling the audience at London's Methodist Central Hall there was an alternative to the "austerity myth" and that Plaid Cymru MPs would work with other "progressive" parties to bring it about.
He was followed by Nicola Sturgeon, who said the SNP wanted to be a voice for Scotland but also a "new progressive politics at Westminster to benefit everyone" in the UK, by "working with other like-minded parties to bring about an alternative to austerity".
The Green Party's Natalie Bennett was the last to give an opening statement, telling the audience the Green Party were the "real challengers" offering an alternative to austerity and fighting against fracking.
There will also be a special Question Time on BBC One, a week before polling day, with Mr Cameron, Mr Miliband and Mr Clegg appearing separately on the programme to answer questions from a studio audience.
Mr Clegg said he had been "denied the opportunity" to speak in Friday's debate because David Cameron had not wanted to take part. Mr Cameron agreed to do one TV debate with six other party leaders, which took place last month.
It was decided that this debate would be for party leaders that were not in government, meaning Mr Clegg was not offered a place on the podium.
Policy guide: Key priorities
What are the top issues for each political party at the 2015 general election?
Responding to Mr Clegg's complaint, a BBC spokesman said broadcasters had worked hard to ensure "that their audiences were offered the best possible combination of programmes to help them engage with the election, to inform them about the issues and to scrutinise the politicians".
He added that the BBC was "satisfied" that its range of election programmes met the obligation to provide "due impartiality" and all relevant parties had the chance to put their case.
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